Despite rumours the couple had started to discuss engagement, the couple have now announced the relationship is over - with the Duchess of Cambridge’s brother apparently the one to call it quits.
A source told The Sun: ‘The split happened a couple of weeks ago and it was James’ decision.’
‘He just didn’t think it was going to go the distance. And I think his family were always very anti her being the future Queen of England’s sister in law.’
Donna, who at 36 is eight years James’ senior, rose to fame when she presented Big Breakfast in the 90s.
And the star apparently confirmed the news by posting a snap form her yoga class, with the quote: ‘Accept what is, let go of what was, have faith in what will be.’
Embarrassingly for Donna, an interview due out this weekend sees her gushing about her royal boyfriend - suggesting that the split has come as a complete surprise to the model.
‘She’s more than a little heartbroken’ a source close to Air told The Sun.
Donna and James met in 2013 through mutual friend Princess Eugenie. After a string of dates they became an item, jetting off on luxurious holidays together (including to the Caribbean with the Middletons just a month ago.)
But despite being pictured looking happy and loved up this summer, it seems things are over for this handsome pair.